In 2017, after doubling the size of the mesocosm array compared to what was used in 2016, we conducted a 16-week experiment that tested the effects of fish and mussels on nutrient recycling and organic matter processing. This experiment addressed several questions: 1) How do green food web responses differ in low flow years under conditions of hydrologic disconnection compared to the same responses in high flow years (as tested by our 2016 mesocosm experiment)? 2) What are the effects of overlapping fish and mussel consumer hotspots on brown (heterotrophic) and green (autotrophic) food webs? The design crossed three levels of fish and three levels of mussel biomass (including a zero-animal mass control), each replicated five times (9 treatments, 45 mesocosms). One manuscript from this experiment has been published (Parr et al., Freshwater Biology, 2019) and the data and analyses are archived here. An additional manuscript is in preparation, and those data will be archived here upon publication.