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Description: The predominant focus in attachment research on trait-like individual differences has overshadowed investigation of the ways in which working models of attachment represent dynamic, interpersonally responsive socio-affective systems. Intensive longitudinal designs extend previous work by evaluating to what extent attachment varies over social interactions and the functional processes that underlie its fluctuation. We examined momentary activation of attachment orientations in the stream of peoples’ daily lives and how those patterns were linked to interpersonal behavior. Based on an event-contingent, ambulatory 7-day assessment protocol (N=263; 3971 interactions) operationalized using Contemporary Integrative Interpersonal Theory, we examined whether contextually activated working models accounted for patterns of interpersonal (anti-)complementarity. Our analyses revealed that the situational activation of working models varied as a function of interpersonal perceptions of warmth, which were linked to greater state security and lower levels of anxious or avoidant expectations. These reactivity patterns, in turn, accounted for interpersonal complementarity. Avoidant attachment was linked to diminished and secure attachment to enhanced expressions of warmth. The analyses were robust even when controlling for momentary positive and negative affect and closeness of the relationship. Attachment expectations wax and wane across daily social interactions, and such fluctuations are reflective primarily of a process in which perceptions of others’ warmth activate secure attachment expectations and lower insecure ones.